Peer-reviewed articles (* high school/undergraduate author)


  1. Haelewaters D, Rossi W. New Laboulbeniales (Fungi, Ascomycota) parasitic on American small carrion beetles (Coleoptera, Leiodidae, Cholevinae). Mycologia.
  2. Pfliegler WP, Báthori F, Wang TW*, Tartally A, Haelewaters D. Herpomyces spp. parasitic fungi (Ascomycota: Laboulbeniales) are globally distributed by invasive cockroach (Insecta: Blattodea) hosts and the pet trade industry. Mycologia.


  1. Haelewaters D, De Kesel A. De schimmel Hesperomyces virescens, een natuurlijke vijand van lieveheersbeestjes. Entomologische Berichten 77 (3): 106-118.
  2. Haelewaters D, Verhaeghen SJC, Ríos González TA, Bernal Vega JA, Villarreal Saucedo RV. New and interesting Laboulbeniales from Panama and neighboring areas. Nova Hedwigia, doi:10.1127/nova_hedwigia/2017/0410 (abstract)
  3. Haelewaters D, Zhao SY, Clusella-Trullas S, Cottrell TE, De Kesel A, Fiedler L, Herz A, Hesketh H, Hui C, Kleespies RG, Losey JE, Minnaar IA, Murray KM, Nedvěd O, Pfliegler WP, Raak-van den Berg CL, Riddick EW, Shapiro-Ilan DI, Smyth RR, Steenberg T, van Wielink PS, Viglášová S, Zhao Z, Ceryngier P, Roy HE. 2017. Parasites of Harmonia axyridis: current research and perspectives. BioControl, doi:10.1007/s10526-016-9766-8 (full text)
  4. Haelewaters D, Pfliegler WP, Szentiványi T, Földvári M, Sándor AD, Barti L, Camacho JJ, Gort G, Estók P, Hiller T, Dick CW, Pfister DH. Parasites of parasites of bats: Laboulbeniales (Fungi: Ascomycota) on bat flies (Diptera: Nycteribiidae) in central Europe. Parasites & Vectors 10: 96. (full text)


  1. Pfliegler WP, Báthori F, Haelewaters D, Tartally A. 2016. Studies of Laboulbeniales on Myrmica ants (III): myrmecophilous arthropods as alternative hosts of Rickia wasmannii. Parasite 23: 50. (full text)
  2. Wang TW*, Haelewaters D, Pfister DH. 2016. Long-hidden in Thaxter’s treasure trove: Laboulbenia camerunensis sp. nov. parasitic on African Curculionidae. Mycotaxon 131 (3): 613-619. (pdf)
  3. Rossi W, Haelewaters D, Pfister DH. 2016. Fireworks under the microscope: a new species of Zodiomyces from Roland Thaxter’s collection. Mycologia 108 (4): 709-715. (full text)
  4. Wang TW*, De Kesel A, Haelewaters D, Pfister DH. 2016. Farlow Herbarium cockroach hosts new record of Laboulbeniales for North America. Rhodora 118 (973): 26-31. (pdf)
  5. Roy HE, Brown PMJ, Adriaens T, Berkvens N, Borges I, Clusella-Trullas S, Comont RF, De Clercq P, Eschen R, Estoup A, Evans EW, Facon B, Gardiner MM, Gil A, Grez AA, Guillemaud T, Haelewaters D, Herz A, Honek A, Howe AG, Hui C, Hutchison WD, Kenis M, Koch RL, Kulfan J, Lawson Handley L, Lombaert E, Loomans A, Losey J, Lukashuk AO, Maes D, Magro A, Murray KM, San Martin G, Martinkova Z, Minnaar I, Nedved O, Orlova-Bienkowskaja MJ, Rabitsch W, Ravn HP, Rondoni G, Rorke SL, Ryndevich SK, Saethre M-G, Sloggett JJ, Soares AO, Stals R, Tinsley MC, Vandereycken A, van Wielink P, Viglášová S, Zach P, Zaviezo T, Zhao Z. 2016. The harlequin ladybird, Harmonia axyridis: global perspectives on invasion history and ecology. Biological Invasions 18 (4): 997-1044. (pdf)
  6. Haelewaters D, Minnaar IA, Clusella-Trullas S. 2016. First finding of the parasitic fungus Hesperomyces virescens (Laboulbeniales) on native and invasive ladybirds (Coleoptera: Coccinellidae) in South Africa. Parasite 23: 5. (full text)


  1. Haelewaters D, Gorczak M, Pfliegler WP, Tartally A, Tischer M, Wrzosek M, Pfister DH. 2015. Bringing Laboulbeniales into the 21st century: enhanced techniques for extraction and PCR amplification of DNA from minute ectoparasitic fungi. IMA Fungus 6 (2): 363-372. (pdf)
  2. Haelewaters D, Boer P, Gort G, Noordijk J. 2015. Studies of Laboulbeniales (Fungi, Ascomycota) on Myrmica ants (II): variation of infection by Rickia wasmannii over habitats and time. Animal Biology 65 (3-4): 219-231. (pdf)
  3. Haelewaters D, Zhao SY, De Kesel A, Royer IR*, Handlin RE*, Farrell BD, Pfister DH. 2015. Laboulbeniales (Ascomycota) of the Boston Harbor Islands I: species parasitizing Coccinellidae and Staphylinidae, with comments on typification. Northeastern Naturalist 22 (3): 459-477. (pdf)
  4. Haelewaters D, Boer P, Noordijk J. 2015. Studies of Laboulbeniales (Fungi: Ascomycota) on Myrmica ants: Rickia wasmannii in the Netherlands. Journal of Hymenoptera Research 44: 39-47. (full text)
  5. Haelewaters D, Rossi W. 2015. Three new species of Laboulbenia from Roland Thaxter’s backlog of slides and a brief review of Laboulbeniales associated with Chrysomelidae. Mycologia 107 (1): 142-148. (full text)
  6. Haelewaters D, Vanpoucke S, Raes D, Krawczynski R. 2015. On carrion-associated beetles in the Sonian Forest (Belgium): observations on five deer carcasses. Bulletin van de Koninklijke Belgische Vereniging voor Entomologie 151: 25-33. (pdf)


  1. Haelewaters D, Yaakop S. 2014. New and interesting Laboulbeniales from southern and southeastern Asia. Mycotaxon 129 (2): 439-454. (pdf)
  2. De Kesel A, Haelewaters D. 2014. Laboulbenia slackensis and L. littoralis sp. nov. (Ascomycota, Laboulbeniales), two sibling species as a result of ecological speciation. Mycologia 106 (3): 407-414. (full text)
  3. Haelewaters D, Vorst O, De Kesel A. 2014. New and interesting Laboulbeniales (Fungi, Ascomycota) from the Netherlands. Nova Hedwigia 98 (1-2): 113-125. (pdf)
  4. Haelewaters D, Comont RF, Zhao SY, Pfister DH. 2014. Hesperomyces virescens (Fungi, Ascomycota, Laboulbeniales) attacking Harmonia axyridis (Coleoptera, Coccinellidae) in its native range. Chinese Science Bulletin 59 (5-6): 528-532. (pdf)
  5. Raak-van den Berg CL, van Wielink PS, de Jong PW, Gort G, Haelewaters D, Helder J, Karssen G, van Lenteren JC. 2014. Invasive alien species under attack: natural enemies of Harmonia axyridis in the Netherlands. BioControl 59 (2): 229-240. (pdf)
  6. Gu X, Haelewaters D, Krawczynski R, Vanpoucke S, Wagner H-G, Wiegleb G. 2014. Carcass ecology: more than just beetles. Entomologische Berichten 74 (1-2): 68-74. (pdf)
  7. Haelewaters D, Schilthuizen M, Pfister DH. 2014. On Diphymyces (Laboulbeniales, Ascomycota) in Malaysian Borneo. Plant Ecology and Evolution 147 (1): 93-100. (pdf)


  1. Haelewaters D. 2013. First record of the genus Ilyomyces for North America, parasitizing Stenus clavicornis. Bulletin of Insectology 66 (2): 269-272. (pdf)
  2. Haelewaters D, De Kesel A. 2013. A new species of Cantharomyces (Laboulbeniales, Ascomycota) from the Netherlands. Mycotaxon 123: 467-472. (pdf)


  1. Haelewaters D, van Wielink P, van Zuijlen JW, Verbeken A, De Kesel A. 2012. New records of Laboulbeniales (Fungi, Ascomycota) for The Netherlands. Entomologische Berichten 72 (3): 175-183. (pdf)
  2. Haelewaters D, Nuytinck J, De Kesel A. 2012. Laboulbeniales (Fungi, Ascomycota) in the Netherlands: an introduction [in Dutch]. Natuurhistorisch Maandblad 101 (5): 88-93. (pdf)

Articles in preparation

  1. Haelewaters D, Howe AG, Meyling NV, Ravn HP, Roy HE, Tragust S, Ceryngier P. Pathogens of the globally invasive Harmonia axyridis (Coleoptera: Coccinellidae): current status and perspectives.
  2. Haelewaters D, De Kesel A. New records of Laboulbeniales (Fungi, Ascomycota) from Canadian insect collection. Nova Hedwigia.
  3. Haelewaters D, De Kesel A, Pfister D.H. Host-associated diversity in a microscopic ectoparasitic fungus.
  4. Haelewaters D, Kappler LA*, Dirks AC*, Buck B, Pfister DH. A Fungal Inventory at the Boston Harbor Islands. Northeastern Naturalist.
  5. Haelewaters D, Gorczak M, Gort G, Pfister DH. Laboulbeniales (Ascomycota) of the Boston Harbor Islands II: species parasitizing Carabidae, including a preliminary morphometric analysis of Laboulbenia flagellata. Northeastern Naturalist.
  6. Haelewaters D, Yaakop S, Rossi W. New and interesting Laboulbeniales from Malaysia.
  7. Haelewaters D. A new species of Diphymyces (Laboulbeniales) from De Kaaistoep. Sterbeeckia.
  8. Gorczak M, Haelewaters D. New species of Laboulbeniales on unusual hosts. Mycological Progress
  9. Haelewaters D, Gorczak M, Pfliegler WP, Pfister DH. The genus Herpomyces (Laboulbeniomycetes).
  10. Ceryngier P, Nedvěd O, Grez AA, Riddick EW, Roy HE, San Martin G, Steenberg T, Veselý P, Zaviezo T, Zúñiga-Reinoso Á, Haelewaters D. Predators and parasitoids of the harlequin ladybird, Harmonia axyridis, in its native range and invaded areas. Biological Invasions.
  11. Saba M, Haelewaters D, Iturriaga T, Khalid AN, Ashraf T, Pfister DH. Geopora ahmadii sp. nov. (Pyrenomataceae, Pezizales), a new species from Pakistan. Mycological Progress.

Miscellaneous publications, not peer-reviewed

  1. Haelewaters D. 2016. Parasietschimmels op mieren. Forum Formicidarum 17 (1): 9-15.
  2. Haelewaters D, van Wielink P. 2016. Hesperomyces virescens (Laboulbeniales) op een nieuwe gastheer. In: Peeters T, van Eck A, Cramer T (eds.) Natuurstudie in De Kaaistoep. Verslag 2015, 21e onderzoeksjaar. TWM Gronden BV, Natuurmuseum Brabant & KNNV-afdeling Tilburg: 9-12.
  3. De Kesel A, Haelewaters D, Dekoninck W. 2016. Myrmecophilous Laboulbeniales (Ascomycota) in Belgium. Sterbeeckia 34: 3-6.
  4. Baral H-O, Haelewaters D. 2015. Rommelaarsia flavovirens gen. et sp. nov. (Helotiales), a new discomycete on Equisetum with a peculiar asexual state. 7 (6) : 321-330.
  5. Roy K., Mann D., Roy H., Murray K. & Haelewaters D. 2015. Museum collections, a treasure trove for fungal hunters. Antenna 39 (4): 178-180.
  6. Haelewaters D, De Kock G, van Wielink P. 2015. Nieuwe Laboulbeniales in De Kaaistoep. In: Peeters T, van Eck A, Cramer T (eds.) Natuurstudie in De Kaaistoep. Verslag 2014, 20e onderzoeksjaar. TWM Gronden BV, Natuurmuseum Brabant & KNNV-afdeling Tilburg: 11-18.
  7. De Kesel A, Haelewaters D. 2014. Belgian records of Laboulbeniales from aquatic insects (3) – species from Dryops luridus. Sterbeeckia 33: 9-15.
  8. De Kesel A, Haelewaters D, Gerstmans C. 2013. Two interesting species of Rickia (Laboulbeniales) from coastal habitats in Belgium and the Netherlands. Sterbeeckia 32: 6-10.
  9. Haelewaters D. 2012. The first record of Laboulbeniales (Fungi, Ascomycota) on ants (Hymenoptera, Formicidae) in The Netherlands. 4 (3): 65-69.

Taxonomic novelties

Genera (1)

Rommelaarsia Baral & Haelew. (2015)

Species (13)

Bordea denotata Haelew. & De Kesel (2014); Cantharomyces elongatus Haelew. & De Kesel (2012); Diphymyces sabahensis Haelew. & Pfister (2014); Gloeandromyces pageanus Haelew. (2017); Laboulbenia bilobata Haelew. & W. Rossi (2015); Laboulbenia camerunensis T.W. Wang & Haelew. (2016); Laboulbenia coccinellidicola Haelew. (2017); Laboulbenia erotylidarum Haelew. (2014); Laboulbenia littoralis De Kesel & Haelew. (2014); Laboulbenia longipilis Haelew. & W. Rossi (2015); Laboulbenia pfisteri Haelew. & W. Rossi (2015); Laboulbenia poplitea Haelew. (2014); Zodiomyces rhizophorus W. Rossi, Haelew. & Pfister (2016).